Zoology - PSEUDOCOELOMATE ANIMALS – Phylum Aschelminthes

 

Things you need to know from this chapter.

 

  1. Definition of a pseudocoelom.
    1. Why isn’t a true coelom?
    2. What are the advantages of a pseudocoelom?
  2. Definition of Aschelminthes
  3. The Aschelminthes that do not molt
    1. Phylum Rotifera

                                          i.    What does a rotifer look like?

                                        ii.    Where do you find rotifers?

                                       iii.    How do rotifers reproduce?

    1. Phylum Acanthocephala

                                          i.    What are they known as?

                                        ii.    What do they prey on?

  1. The Aschelminthes that do molt
    1. Phylum Nematoda

                                          i.    General characteristics of all roundworms.

                                        ii.    Know the general life cycles & ways to avoid infection with the

                                       iii.    following parasites…

1.    Ascaris lumbricoides

2.    Necator americanus

3.    Trichinella spiralis

4.    Filarial worms

                                       iv.    What causes elephantiasis?  How is it spread?

                                        v.    What causes river blindness?  How is it spread?

                                       vi.    What causes dog heart worms?  How are they spread?

    1. Phylum Nematomorpha

                                          i.    Why are they called horsehair worms?

                                        ii.    Know their general life cycle.

    1. Phylum Kinorhyncha

                                          i.    Where do you find them?

                                        ii.    What do they eat?

                                       iii.    How do they reproduce?

    1. Phylum Loricifera

                                          i.    When were they discovered?

                                        ii.    Where do you find them?

    1. Phylum Priapulida

                                          i.    Where do you find them?

                                        ii.    How do they eat?

 

I.  Compared and Contrasted to Acoelomate Animals

 

            A.  They are like the acoelomate animals in that they…

                        1.  They are bilateral “worms”

                        2.  Cephalization

                        3.  Lack circulatory and respiratory systems

 

            B.  They are more complex than the acoelomate animals in that they…

                        1.  They have a pseudocoelom derived from the blastocoel.

                        2.  They are more mobile.

                        3.  They have more complex reproductive systems.

                        4.  They can store wastes for discharge out of the body.

                        5.  They have a complete mouth to anus digestive system

 

II. What is a pseudocoelom?

 

A. Structure

1. The original ______________________ of the embryo persists

as a space or cavity between digestive cavity and the body wall.

2. Since it lacks the peritoneal lining of a true coelom, it is a

Pseudocoel “____________________________________”.

 

B. Biological Contributions

1. Pseudocoelom generally provides greater freedom of movement.

2. There is space for more development of digestive, excretory, and

reproductive systems.

3. It is simpler to distribute materials throughout the body.

4. This provides a storage area for waste products to be later

discharged.

5. As a ______________________ organ, the cavity is a base for

muscles that provide movement.

 

III. Pseudocoelomates

 

A.    A term which refers to animals with a pseudocoelomate body plan is Aschelminthes.

1.       _____________________________________ -- A heterogeneous phylum of small to microscopic wormlike animals; individuals are pseudocoelomate and

mostly unsegmented and are covered with a cuticle.

 

B.  Except in parasitic ______________________, the digestive tract is

complete (from mouth to pharynx to intestines to anus).

 

C. Digestive tract, gonads and excretory organs are within the

______________________.

 

D. In some, the epidermis secretes a nonliving

______________________.

 

IV.  Lophotrochozoan phyla – the Aschelminthes that do NOT molt

 

A.    Phylum Rotifera

 

1. Rotifers have a ciliated crown, the ______________________ that

beats like a rotating wheel.

a. Rotifers are highly diverse in color, size and shape; some are

colonial.

b. Rotifers are found worldwide (______________________

distribution) with over 1,800 species known.

c. Most are freshwater species but a few are

______________________ (living in salt water),

______________________ (living on dry land) or parasitic.

d. Many endure ______________________ (drying out) and

temperature changes by ______________________.

 

2. Reproduction

a. Rotifers are usually ______________________

1) Males are smaller than females.

2) In some classes, males are unknown, and in others,

males occur only briefly.

b. Bdelloidea females are ____________________________,

producing diploid eggs that hatch into diploid females.

c. Monogononta females produce diploid

______________________ eggs that form diploid females, or

haploid ______________________ eggs that, if not fertilized,

become haploid males.

 

B.    Phylum Acanthocephala _____________________________ worms

 

1. All spiny-headed worms are parasites in the intestines of

______________________.

a. Its proboscis has rows of recurved spines that penetrate and

may rupture host intestines.

b. Larvae develop in crustaceans or insects.

c.  They never infest humans.

 

2. Form and Function

a. The body is somewhat flattened.

b. Both longitudinal and circular body wall muscles are present.

c. There is no respiratory system or heart.

d. They lack a digestive tract and absorb all nutrients across the

tegument, which bears some enzymes.

e. Shelled embryos discharged in the feces do not hatch until

eaten by an intermediate host, often _____________________.

f. Larval acanthors burrow through beetle intestine and develop

into juvenile cystacanths in the insect hemocoel.

 

V.  The Ecdysozoan phyla – Aschelminthes that do molt

 

A.    Phylum Nematoda: ______________________

 

1. About 12,000 species are described; perhaps a half million exist.

a. They live in virtually all habitats in all biomes; topsoil may

contain billions per acre.

b. Nematode parasites exist in nearly all animal and plant

species;

            1)  They are economically important.

c. Free-living nematodes feed on bacteria, yeasts, fungal

hyphae and algae.

d. Predatory nematodes eat rotifers, tardigrades, small

annelids and other nematodes.

e. Nematodes are also important as food for mites, insects,

larvae and fungi.

 

2. Form and Function

a. Distinguishing Characteristics

1) They have a cylindrical shape.

2) They have nonliving cuticle.

3) Muscles in the body wall only run longitudinally.

b. The pseudocoel serves as a hydrostatic skeleton against

which longitudinal muscles work.

c. An outer, thick, noncellular cuticle is secreted by the

underlying hypodermis.

d. The cuticle has layers of crisscrossing collagen, providing

elasticity but constraining expansion.

e. Digestion

1) The alimentary canal consists of mouth, pharynx, a

non-muscular intestine, a short rectum and the anus.

                        f.  Reproduction

1) Most nematodes are dioecious with males smaller

than females.

            2)  The male has copulatory spicules to hold the

            female vulva open against hydrostatic pressure.

3) Fertilization is internal and eggs are stored in the

uterus until deposited.

 

3. Some nematodes are important pathogens of humans; most are tropical.

a. The ________________________________________of

humans: Ascaris lumbricoides

1) Ascaris lumbricoides occurs in up to 64% of people

in some areas of the southeastern U.S.

2) More than 1.2 billion are affected worldwide.

3) A female Ascaris lays 200,000 eggs a day, passing

out in the host’s ______________________.

4) Embryonic development completes in two weeks.

5) Viable eggs remain after signs of fecal matter have

disappeared; eggs survive long periods in soil.

6) When a host swallows ______________________,

juveniles hatch and burrow through intestinal wall.

7) Carried through the heart to the

______________________, they break into the

alveoli and are carried up to the tracheae.

8) Coughed up and swallowed, they mature in the

______________________ two months after they were swallowed.

9) They feed on ______________________________

and may block or perforate the intestines.

10)  Infection avoidance: 

 

 

 

b. Enterobius vermicularis: Pinworms

1) It is the most common ______________________

parasite in the U.S. but causes little disease.

2) Adults live in the large intestine and cecum.

3) Females, about 12 mm long, migrate to the anal

region at night and lay eggs, causing ____________.

4) Scratching the anal region contaminates hands and

bedclothes.

5) Eggs develop rapidly and become infective within

six hours at body temperature.

6) When ______________________, they hatch in

the duodenum and mature in the large intestine.

7) Members of this order have haploid males from

unfertilized eggs; females are diploid and come from

fertilized eggs.

8)  Infection avoidance: 

 

 

 

 

 

 

c. The most common human ______________________:

Necator americanus

1) The anterior end of these small (9-11 mm) worms

has a hook-like curve.

2) The males and females are separate.

3) Large plates in their mouths cut into intestinal

mucosa; then they suck the ___________________.

4) They pump through more blood than they digest;

heavy infections cause ______________________.

5) Eggs pass in ______________________ and

juveniles hatch in soil where they live on

______________________.

6) If human skin comes in contact with soil, infective

juveniles burrow through ______________________

to ______________________.

7) Similar to Ascaris, they travel in blood to lungs, are

coughed up to be swallowed, and mature in the

intestine.

8)  Infection avoidance: 

 

 

 

d. Trichina Worm:  ________________________________

1) While tiny, this worm can cause potentially lethal

______________________.

2) Adult worms burrow into the intestinal mucosa and

females directly produce ______________________.

3) Juveniles penetrate blood vessels and circulate

throughout the body to all tissues and spaces.

4) They penetrate skeletal muscle cells, redirecting

gene expression of the musculature so it loses its

striations and becomes a _____________________to

the parasite.

5) When meat containing live juveniles is eaten,

 worms are liberated and mature in the

______________________.

6) They infect humans, hogs, rats, cats and dogs;

hogs can become infected eating uncooked scraps of

infected meat or rats.

7) Heavy infections cause death; about 2.4% of the

U.S. population is infected, mostly lightly.

8)  Infection avoidance: 

 

 

 

5. Filarial Worms - Eight species of filarial nematodes infect

humans; some cause major and serious diseases. 

a. Wucheria bancrofti and Brugia malayi live in the

______________________ system.

1) The worms cause inflammation and

blockage of the lymphatics.

2) Females release live young, tiny

______________________, into blood and

lymph.

3) ______________________ ingest the

microfilariae when they feed; the worms

develop to the infective stage and move into

the mosquito bite wound when it feeds.

4) ______________________ is caused by

repeated exposure; swelling and growth of

connective tissue causes enormous swelling of

body parts.

b. River blindness or ______________________ is

carried by ______________________ and infects 30 million people in tropics.

c. Dog heartworm, ___________________________,

is carried by mosquitoes and is most common U.S.

filarial worm.

 

B. Phylum Nematomorpha – “________________________________

 

1. Resemble coarse hairs from a horse’s tail.

            a.  People believed they spontaneously generated when a

            horse’s tail fell into water.  The name stuck.

 

2. Form and Function

a. Adults are free-living in moist habitats; juveniles are

parasites of ______________________.

b. They range from 10 to 70 cm long but only 0.3 to 2.5 mm

in diameter.

c. Circulatory, respiratory and excretory systems are lacking.

d. Juveniles only emerge from the arthropod host when

______________________ is nearby.

e. Females discharge eggs into water; juveniles hatch and

gain entry to the arthropod host.

f. After months in an arthropod host, the mature worm

emerges into nearby water or during rainfall.

g. Somehow, the parasite stimulates terrestrial insects to

seek water.

 

C. Phylum Kinorhyncha

 

1. Kinorhynchs are small (usually less than 1mm long) marine

animals that feed on ______________________ (a photosynthetic

single celled organism enclosed by a shell of silica).

a. About 75 species are known.

b. They are cosmopolitan.

c. The body is divided into 13 segments with spines but no

cilia.

d. It cannot swim, but anchors in its silt or mud burrow with

spines.

            2.  Reproduction

a. Sexes are separate with paired gonads and gonoducts.

b. Development includes a series of six juvenile stages and

a nonmolting adult.

 

D. Phylum Loricifera

 

1. Loriciferans were recently discovered (____________________) in spaces between marine grains.

a.  They are tiny, less than ¼ mm.

b.  They are apparently widely distributed.

 

E. Phylum Priapulida

 

1. 18 species of these marine worms occur in colder waters.

a. Some are tube dwellers and feed on _________________

(a mass of dead or decaying organic matter).

b. Priapulids have cylindrical bodies under 15 cm long.

c. They burrow by body contractions and orient their mouth

at the surface.

d. A chitinous cuticle covers the body and is molted

regularly.