Zoology – ORGANIC EVOLUTION

 

I. Origins of Darwinian Evolutionary Theory

 

A. Pre-Darwinian Evolutionary Ideas

1. Before the 18th century, speculation on origin of species was not

scientific.

2. _________________________________ portrayed a constant

world after a creation event.

3. Early Greek philosophers considered some ideas of evolutionary

change.

a. _____________________ were recognized as former life

destroyed by _________________________________.

c. Lacking a full evolutionary concept, the idea faded before

the rise of _____________________.

4. Biblical account of creation became a tenet of faith.

a. Evolutionary views were heretical.

b. __________________________________________

calculated _____________________ as date of life’s

creation.

5. French biologist Jean Baptiste de _____________________

offered first complete explanation in _____________________.

a. He convincingly argued that fossils were remains of

_____________________ animals.

b. Lamarck’s mechanism was inheritance of ___________

_____________________.

c. He explained long necks of giraffes to stretching efforts of

ancestral giraffes.

d. Lamarck’s concept is ____________________________;

individuals transform their own traits to evolve.

e. In contrast, Darwin’s theory is ______________________

or due to differential survival among offspring.

7. Geologist Sir Charles _____________________ established the

principle of _____________________.

a. Uniformitarianism consists of two important principles:

1) Laws of physics and chemistry remain the same

throughout earth’s history.

2) Past geological events occurred by natural

processes similar to those observed today.

b. Natural forces acting over long periods could explain

formation of fossil-bearing rocks.

c. Earth’s age must be measured in

_____________________ of years.

          1)  It is now accepted that earth is

          _____________________ years old.

          2)  Life has existed on earth for more than 3.5 billion

          years.

d. Geological changes are natural and without direction; both

concepts underpinned Darwin.

 

B. Darwin’s Great Voyage of Discovery (_____________________)

1. In 1831, Charles Darwin (almost 23) sailed aboard the small

survey ship _____________________.

2. Darwin made extensive observations in the five-year voyage.

a. Darwin collected the fauna and flora of _______________

_____________________ and adjacent regions.

b. He unearthed long extinct fossils and associated fossils of

South and North America.

c. He saw fossil seashells embedded in the Andes rocks at

13,000 feet altitude.

d. Observing earthquakes and severe erosion confirmed his

views of geological ages.

3. The __________________________________________

provided unique observations.

a. These volcanic islands are on the equator 600 miles west

of _____________________.

b. Each island varied in tortoises, iguanas, mockingbirds and

ground finches.

c. The islands had similar climate but varied vegetation.

d. Island species therefore originated from South America

and were modified under the varying conditions of different

islands.

4. Darwin conducted the remainder of his work at home in

_____________________.

                    a. In 1838, Darwin read an essay on population by Thomas

                    R. _____________________.

b. Having studied artificial selection, a “struggle for

existence” because of overpopulation gave him a

mechanism for evolution of wild species by natural

selection.

c. In 1858, he received a manuscript from a young naturalist,

__________________________________________,

summarizing the main points of natural selection.

d. Geologist Lyell and botanist Hooker persuaded Darwin to

publish a paper jointly with Wallace’s paper.

e. Darwin then rushed to publish a shorter “abstract” version

in 1859: ________________________________________

_______________________________________________.

f. 1250 copies sold of first printing in one day.

 

SHOW THE GREAT BOOKS SERIES:  THE ORIGIN OF SPECIES

 

II.  Darwin’s Theory – Evolution Occurs by _____________________________

 

A. Natural selection gives a natural explanation for origins of adaptation.

 

B. Darwin’s theory of natural selection consists of four observations and

three conclusions.

1. Observation 1:  _____________________________________

_____________________________________________________.

a) If all individuals produced survived, populations would

explode exponentially.

b) Darwin calculated that a single pair of elephants could

produce 19 million offspring in 750 years.  This is a SLOW

breeding species!

1)  Elephant females breed at the earliest at 9 years,

and bear one calf every four years until age fifty

optimally.  It would take a female 40 years to bear 10

calves.

2. Observation 2:  _____________________________________

_____________________________________________________

_____________________________________________________

_____________________________________________________.

a)  Conclusion 1:  The struggle for food, shelter, and space

becomes increasingly severe with overpopulation.  There is

competition for __________________________________

______________________________________________.

          1)  In each generation, many individuals must die

          young, fail to reproduce, produce few offspring, or

          produce less-fit offspring that fail to survive &

          reproduce in their turn.

          2)  Survivors represent only a small part of those

          produced each generation.

3. Observation 3:  _____________________________________

__________________________________________.  Individual

members of a population differ from one another in their ability to

obtain resources, withstand environmental extremes, escape

predators, etc.

          a)  Conclusion 2:  The most well-adapted (the “fittest”)

          individuals in one generation will usually leave the most

          offspring.  This is _________________________________: 

          the process by which the environment selects for those

          individuals whose traits best adapt them to that particular

          environment.

4.  Observation 4:  _____________________________________

_____________________________________________________

_____________________________________________________.

a) Darwin only noted the resemblance of parents and

offspring.

b) Gregor Mendel’s mechanisms of heredity were applied to

evolution many years later.

c)  Conclusion 3:  Over many generations, differential, or

unequal, reproduction among individuals with different

genetic makeup changes the overall genetic composition of

the population.  This generates new adaptations and new

species.

 

C. Natural selection can be viewed as a two-part process: random and

non-random.

1.  Production of variation among organisms is random; mutation

does not generate traits preferentially.

2.  The nonrandom component is the survival of different traits.

 

SHOW BIOVIDEO:  THE EVIDENCE FOR EVOLUTION

 

III. Evidence for Evolution

 

          A.  The fossil record provides evidence of evolutionary change over time.

1. The living world is constantly changing in form and diversity. 

Change in animal life is directly seen in the 600-700 million-year

animal fossil history.

2. A fossil is a remnant of past life.

a. Insects in amber and frozen mammoths are actual

remains.

b. Teeth and bones can petrify or become infiltrated with

silica and other minerals.

c. Molds, casts, impressions and fossil excrement are also

fossils.

3. Most organisms leave no fossils; the record is always incomplete

and requires interpretation.

4. Interpreting the Fossil Record

a. The fossil record is biased because preservation is

selective.

b. Vertebrate skeletons and invertebrates with shells provide

more records.

c. Soft-bodied animals leave fossils only in exceptional

conditions such as the Burgess Shale.

d. Fossils occur in stratified layers; new deposits are on top

of older material.

          1)  The law of stratigraphy dates oldest layers at the

          bottom and youngest at the top.

e. “Index” or “guide” fossils are “indicators” of specific

geological periods.

5. Radiometric Dating

a. In the late 1940s, this dating method was developed that

determines age of rocks.

b. Radioactive decay of naturally occurring elements is

independent of heat and pressure.

c. Potassium-Argon Dating

1) Potassium-40 (40K) decays to argon-40 (40Ar) and

Calcium-40 (40Ca).

2) Half-life of potassium-40 is 1.3 billion years; half of

remainder will be gone at end of next 1.3 billion years,

etc.

3) Calculating the ratio of remaining potassium-40 to

amount originally there provides mathematically close

estimate of age of deposit.

6. Evolutionary Trends - Fossil record allows observation of

evolutionary change over broad periods of time.

a. Animals species arise and go extinct repeatedly.

b. Animal species typically survive 1-10 million years; there

is much variability.

c. Horse Evolution Shows Clear Trend

1) From Eocene to Recent periods, genera and

species of horses were replaced.

2) Earlier horses had smaller sized and fewer grinding

teeth, and more toes.

3) Reduction in toes and increase in size and

numbers of grinding teeth correlate with

environmental changes.

 

B. Common Descent – Darwin proposed that all plants and animals

descended from a common ancestor. 

          1.  Geographic Animal Distribution and continental drift

                    a.  Large flightless birds exist only in the southern

                    hemisphere

                              1) extant rhea of South America & emu of Australia

                              2) extinct giant moa of New Zealand & elephant bird

                              of Madagascar

                    b.  These birds are flightless, their ancestor couldn’t have

                    swum the oceans to these continents.  The explanation is

                    that their ancestor evolved on the southern continent of

                    Gondwanaland.  (The northern continent was Laurasia.)

          2.  Comparative anatomy

                    a.  All plants and animals descending from a common

                    ancestor is divergent evolution.

                              1)  Homologous structures – structures that may

                              differ in function but that have similar anatomy due to

                              descent from a common ancestor.

                                        a)  For example, vertebrate limbs show the

                                        same basic structures (one upper arm bone,

                                        two forelimb bones, wrist bones, and finger

                                        bones) modified for different functions

                                        (swimming in penguins & whales, flight in bats

                                        & birds, running in dogs & sheep, grasping in

                                        humans & shrews)

                                        b)  Darwin saw homology as major evidence

                                        for common descent.

                                        c)  It is inconceivable that nearly the same

                                        bone arrangements could be ideal for such

                                        different functions, as would be expected if

                                        each animal were created separately.  This is

                                        exactly what we would expect if birds &

                                        mammal forelimbs evolved from a common

                                        ancestor.

                              2)  Vestigial structures – structures that serve no

                              apparent purpose

                                        a)  For example, vampire bats have molar

                                        teeth even though they live on a diet of blood &

                                        therefore don’t chew their food.

                                        b)  Pelvic bones exist in whales and certain

                                        snakes.

                                        c)  This is “evolutionary baggage”

                                        d)  The appendix is a vestigial structure in

                                        humans

                    b.  Through convergent evolution, natural selection has

                    shaped unrelated organisms into similar forms in similar

                    environments.

                              1)  Natural selection predicts that, given similar

                              environmental demands, unrelated organisms might

                              independently evolve superficially similar structures.

                              2)  Such outwardly similar body parts in unrelated

                                        organism, called analogous structures, often have

                              completely different internal anatomy because they

                              are not derived from a common ancestor.

                              3)  For example:  wings of flies & birds; also fat-

                              insulated, streamlined shapes of seals (mammals) &

                              penguins (birds)

 

          C.  Modern Biochemical & genetic analyses reveal relatedness among

          diverse organisms

                    1.  The amino acid sequences of proteins are remarkably similar

                    across a huge spectrum of species.

                    2.  The chromosomes of chimpanzees & humans are extremely

                    similar, showing that these species are closely related.

 

          D.  Artificial selection demonstrates that organisms may be modified by

          controlled breeding

                    1.  In a few hundred to at most a few thousand years, man has bred

                    radically different dog breeds, from Chihuahua to Great Dane, from

                    the wolf by selecting qualities that were found desirable &

                    selectively breeding for them.

                    2.  What man can do, cannot nature do by selecting for those

                    individuals best adapted to that environment

 

          E.  Evolution by Natural Selection Occurs Today

                    1.  We can literally see that evolution is occurring around us today

                    2.  Peppered moths of England & the Industrial Revolution

                    3.  1990’s Florida’s cockroaches, Combat, corn syrup with glucose

                    as bait, rare mutation became widespread

                    4.  Any pesticide, herbicide, or antibiotic & resistance